Velarion · Company Intelligence
Compensation Brief
Fiscal year 2025 · Filings through 2026-05-28

aTYR PHARMA (ATYR)

Fiscal year 2025 executive and director compensation, incentive-plan design, equity activity, and governance — every figure in this brief is traceable to its exact location in the company’s SEC filing — the document, the section, the page, and the table, linked directly.

$3,456,253
Sanjay Shukla total, FY2025
33rd
CEO pay percentile vs peers
CEO-to-median pay ratio
97.61%
Say-on-pay support, latest vote
Companies that benchmark against ATYR
4 companies name ATYR as a compensation peer

Each company listed discloses ATYR in its own benchmarking peer group, as disclosed in each company's most recent proxy statement within the last two proxy seasons. Two companies are therefore absent by design: one that named ATYR in an earlier year and no longer does, and one whose own most recent disclosure is older than the last two seasons.

01

Summary Compensation Table — fiscal year 2025

Named executive officerSalaryBonusStock awardsOption awardsNon-equity incentivePension / NQDCAll otherTotal
Sanjay Shukla$638,977$0$0$2,595,200$210,862$0$11,214$3,456,253
President and Chief Executive Officer
Jill Broadfoot471,79000713,680113,229011,2141,309,913
Chief Financial Officer
Nancy Denyes451,57000713,680108,377011,2141,284,841
General Counsel and Corporate Secretary

A dash means the company reported no figure in that column; 0 means it reported zero. Figures are as printed in the filing, without adjustment.

Three-year trend — total compensation per named executive

Named executive officerFY2025 totalFY2024 totalFY2023 totalΔ FY20242025
Sanjay Shukla$3,456,253$1,480,522+1,975,731
Jill Broadfoot1,309,913746,163+563,750
Nancy Denyes1,284,841729,305+555,536

Prior-year totals as disclosed in the corresponding proxy; deltas are computed values with the derivation shown in the register.

02

Equity transactions since the proxy statement was filed

Every insider equity transaction reported on Form 4 after the 2025 proxy statement — acquisitions, dispositions, exercises, and withholdings — and therefore not reflected in any table in that document.

InsiderRoleTransaction dateCodeSharesPriceValueSecuritySource
Svetlana LucasDirector2026-05-11A50,000$0$0Derivative, Class A CommonForm 4 ↗
Sara ZaknoenDirector2026-05-11A50,00000Derivative, Class A CommonForm 4 ↗
Paul SchimmelDirector2026-05-11A50,00000Derivative, Class A CommonForm 4 ↗
Eric BenevichDirector2026-05-11A50,00000Derivative, Class A CommonForm 4 ↗
Timothy P. CoughlinDirector2026-05-11A50,00000Derivative, Class A CommonForm 4 ↗
Jane A. GrossDirector2026-05-11A50,00000Derivative, Class A CommonForm 4 ↗
Paul SchimmelDirector2026-05-27A100,000049,630CommonForm 4 ↗

Transaction codes per Form 4: A acquisition · S open-market sale · F tax withholding · M option exercise · G gift. Value = shares × price as reported. Dividend-equivalent and fractional-share accrual rows (derivative Class A, under $10,000 and under 2,000 shares) are aggregated per date; every row appears in full below.

03

Grants of plan-based awards — fiscal year 2025

ExecutiveAwardGrant dateThresholdTargetGrantedMaximumGrant-date fair value
Sanjay ShuklaOption$2,595,200
Nancy DenyesOption713,680
Jill BroadfootOption713,680
03b

Vehicle mix — grant-date fair value by award type

Named executive officerTime-vested (RSU)Performance-vested (PSU)Options / SARs% performance-vested
Sanjay Shukla$0$0$2,595,2000.0%
Nancy Denyes00713,6800.0%
Jill Broadfoot00713,6800.0%

% performance-vested = PSU grant-date fair value ÷ (RSU + PSU + Options), a computed value with the derivation shown in the register.

04

Outstanding equity awards at fiscal year-end 2025

ExecutiveAwardShares unvestedExercise priceMarket value unvested
Nancy DenyesRSU2,687$2,104
Nancy DenyesOption8965.52
Sanjay ShuklaOption770,8343.74
Sanjay ShuklaRSU10,3758,125
Sanjay ShuklaOption312,5001.5
Jill BroadfootOption7,8492.81
Jill BroadfootOption1,2505.52
Jill BroadfootOption24,5072.2
Jill BroadfootOption211,9803.74
Nancy DenyesOption9,1772.81
Nancy DenyesOption211,9803.74
Nancy DenyesOption78,1251.5
Nancy DenyesOption24,5072.2
Jill BroadfootRSU3,7502,937
Sanjay ShuklaOption3,4585.52
Sanjay ShuklaOption53,7792.81
Sanjay ShuklaOption98,0282.2
Jill BroadfootOption78,1251.5
05

Incentive-plan design — the Compensation Discussion & Analysis

Our executive compensation philosophy is based on the following objectives: Aligning executive interests and stockholder interests through long-term incentives linked to Company performance; Attracting, retaining and motivating superior executive talent who exemplify and enhance the Company culture; and Providing incentives that reward the achievement of performance goals that directly correlate to the enhancement of stockholder value, as well as to facilitate executive retention.

CEO STI target (% of salary)55%
CEO LTI target (% of salary)
Independent consultantRadford (Aon plc Rewards Solutions)
Clawback policyTrue
Anti-hedging
Anti-pledging
06

Say-on-pay — recent advisory votes

Meeting yearMeeting dateForAgainstAbstainSupportOutcome
20252025-05-0155,237,7191,353,536256,53597.61%Passed
20242024-05-2242,468,397397,92650,83399.07%Passed
20232023-05-1737,246,0691,007,21245,49797.37%Passed
20222022-04-2618,771,20251,25432,56399.73%Passed
20212021-04-287,519,078136,22818,01198.22%Passed

Rows are labeled by annual-meeting year, with the meeting date beside each; the vote held in a given year acts on the prior fiscal year's compensation.

Say-on-pay — history, company's computation

Meeting yearMeeting dateApproval — company's computation
20252025-05-0197.61%a
20242024-05-2299.07%b
20232023-05-1797.37%c

a. Approved under the standard stated in the filing — “Broker non-votes and abstentions will not be counted as votes properly cast with respect to such matter, and will have no effect on the proposal.”.

b. Approved under the standard stated in the filing — “Broker non-votes and abstentions will not be counted as votes properly cast with respect to such matter, and will have no effect on the proposal.”.

c. Approved under the standard stated in the filing — “Broker non-votes and abstentions will not be counted as votes properly cast with respect to such matter, and will have no effect on the proposal.”.

Approval percentages are the company's own reported computations under its own stated standard, not recomputed by Velarion. Where a company does not state a standard, none is assumed.

07

CEO pay ratio

Ratio (CEO : median employee)
CEO total compensation
Median employee compensation
08

Director compensation — fiscal year 2025

DirectorCash feesStock awardsOption awardsNon-equity incentiveAll otherTotal
Timothy P. CoughlinIndependent · Board Chair · Audit$82,200$0$112,085$0$194,285
Svetlana Lucas, Ph.DIndependent · Audit (Chair) · Nominating/Governance63,0440112,0850175,129
Jane A. Gross, Ph.DIndependent · Compensation · Nominating/Governance58,3380112,0850170,423
Sara L. Zaknoen, M.DIndependent · Compensation · Nominating/Governance55,3370112,0850167,422
Eric BenevichIndependent · Audit · Compensation51,7640112,0850163,849
Paul Schimmel, Ph.DIndependent · Not standing for re-election40,0000112,0850152,085
John K. ClarkeIndependent · Departed during FY19,7750019,775
Sanjay S. Shukla, M.D., M.SEmployee director00000
09

Board fee structure and ownership guideline

Annual cash retainer
Annual equity retainer
Per board meeting fee
Committee chair fees (audit / comp / nom-gov)20,000 / 15,000 / 10,000
Stock ownership guideline
09b

Total cost of the board

ComponentFY2025
Cash retainers and fees$370,458
Equity awards (grant-date value)$0
All other compensation$0
Total cost of the board$370,458

Computed as the sum of the director compensation table rows above; inputs and formula shown in the register.

10

Disclosed peer group — fiscal year 2025

The compensation peer group as the company disclosed it in its own proxy — the reference set behind the percentile above.

10a

Peer reciprocity

Of the companies aTYR PHARMA names as compensation peers, which name it back in their own disclosed peer groups. A peer set the market reciprocates reads differently from one selected in a single direction.

Reciprocity shown where the peer's own current proxy has been read — each mark is cited in the register to that peer's filing. A dash means the peer's current proxy was read and names no reciprocal group including aTYR PHARMA.

10b

Chief executive pay against the peer group

Most recent fiscal-year chief executive compensation at each disclosed peer, taken from that company's own proxy statement.

RankCompanyFYChief executiveSalaryStock awardsOption awardsNon-equity incentiveTotal
1Scholar Rock Holding Corporation2025David Hallal$637,500$22,518,500$7,624,988$452,293$33,300,927
2Altimmune, Inc.2025Vipin Garg658,6941,153,6004,812,167357,7538,026,590
3KalVista Pharmaceuticals, Inc.2024Benjamin Palleiko580,4415,486,659302,46806,772,197
4Allakos Inc.2023Robert Alexander805,4602,171,3402,623,519684,6416,299,426
5Pliant Therapeutics, Inc.2025Bernard Coulie682,0001,810,2502,713,750204,6005,462,901
6Savara Inc.2025Matthew Pauls660,0003,870,0000297,0004,845,000
7aTYR PHARMA2025Sanjay Shukla638,97702,595,200210,8623,456,253
8X4 Pharmaceuticals, Inc.2025Paula Ragan420,1701,081,575002,640,569
9Gossamer Bio, Inc.2025Faheem Hasnain649,64901,392,006254,8002,300,455
10Longboard Pharmaceuticals, Inc.2023Kevin Lind610,0000986,934442,8602,053,804
11Alector, Inc.2025Arnon Rosenthal678,100422,876558,216325,4881,995,018
Kezar Life Sciences, Inc.not in coverage universe
Immunic, Inc.not in coverage universe
Trevi Therapeutics, Inc.not in coverage universe
Fortress Biotech, Inc.not in coverage universe
Jasper Therapeutics, Inc.not in coverage universe

Each peer's compensation is from its most recently filed proxy; fiscal years differ by company. Each peer's figures come from its own filings; the subject company is highlighted. Peers whose chief executive compensation is outside our coverage universe are stated as such, never left blank.

10c

Incentive design against the same peers

CompanyAnnual incentive payoutLong-term vehicle mixPSU measures include TSRCompensation consultant
aTYR PHARMA60% (Committee lowered payout from 75% achievement to 60% given EFZO-FIT study results)%Options 100not disclosedRadford (Aon plc Rewards Solutions)
Alector, Inc.80%not disclosednot disclosedPearl Meyer & Partners, LLC
Allakos Inc.not disclosedOptions 55 · RSU 45NoCompensia
Altimmune, Inc.97.5%not disclosednot disclosedNational compensation consulting firm (unnamed)
Fortress Biotech, Inc.not disclosednot disclosednot disclosednot disclosed
Gossamer Bio, Inc.65%not disclosedNoAlpine Rewards
Immunic, Inc.not disclosednot disclosednot disclosednot disclosed
Jasper Therapeutics, Inc.not disclosednot disclosednot disclosednot disclosed
KalVista Pharmaceuticals, Inc.not disclosednot disclosedNoAon
Kezar Life Sciences, Inc.not disclosednot disclosednot disclosednot disclosed
Longboard Pharmaceuticals, Inc.not disclosednot disclosednot disclosednot disclosed
Pliant Therapeutics, Inc.50%Options 67 · RSU 33not disclosedAlpine Rewards
Savara Inc.90%RSU 100not disclosedAon
Scholar Rock Holding Corporationnot disclosednot disclosedNoPay Governance
Trevi Therapeutics, Inc.not disclosednot disclosednot disclosednot disclosed
X4 Pharmaceuticals, Inc.not disclosednot disclosednot disclosednot disclosed

Read from each company's Compensation Discussion and Analysis: annual incentive payout as a percentage of target, the disclosed long-term vehicle mix, whether performance-share measures include a total shareholder return component, and the retained compensation consultant. “Not disclosed” appears only where the company's filing does not state the item.

11

Source register

Each figure's location is given both for the eye (section · page · table · row · column) and for a script (table/row/column index and exact character span in the filed document). Any figure can be independently confirmed against the source.

#SupportsDocumentFiledAccessionLocation in document
1Section 01, Sanjay ShuklaDEF 14A2026-03-260001339970-26-000021Section “SUMMARY COMPENSATION TABLE”. As printed: 3,456,253. Filing ↗machine: t11 · r2 · c21 · span 533754–533763
2Section 01, Jill BroadfootDEF 14A2026-03-260001339970-26-000021Section “SUMMARY COMPENSATION TABLE”. As printed: 1,309,913. Filing ↗machine: r4 · c21 · span 549228–549237
3Section 01, Nancy DenyesDEF 14A2026-03-260001339970-26-000021Section “SUMMARY COMPENSATION TABLE”. As printed: 1,284,841. Filing ↗machine: r6 · c21 · span 564707–564716
4Section 03, Sanjay Shukla OptionDEF 14A2026-03-260001339970-26-000021Section “SUMMARY COMPENSATION TABLE” · grants of plan-based awards table. Filing ↗machine: t11 · r2 · c9 · span 529802–529811
5Section 03, Nancy Denyes OptionDEF 14A2026-03-260001339970-26-000021Section “SUMMARY COMPENSATION TABLE” · grants of plan-based awards table. Filing ↗machine: t11 · r6 · c9 · span 560757–560764
6Section 03, Jill Broadfoot OptionDEF 14A2026-03-260001339970-26-000021Section “SUMMARY COMPENSATION TABLE” · grants of plan-based awards table. Filing ↗machine: t11 · r4 · c9 · span 545278–545285
7Section 07, pay ratioDEF 14A2026-03-260001339970-26-000021Section “CEO Pay Ratio”. As printed: —. Filing ↗
8Section 08, Timothy P. CoughlinDEF 14A2026-03-260001339970-26-000021Section “Director Compensation”. As printed: 194,285. Filing ↗machine: t19 · r2 · c11 · span 1219152–1219159
9Section 08, Svetlana Lucas, Ph.DDEF 14A2026-03-260001339970-26-000021Section “Director Compensation”. As printed: 175,129. Filing ↗machine: t19 · r6 · c11 · span 1238999–1239006
10Section 08, Jane A. Gross, Ph.DDEF 14A2026-03-260001339970-26-000021Section “Director Compensation”. As printed: 170,423. Filing ↗machine: t19 · r5 · c11 · span 1234049–1234056
11Section 08, Sara L. Zaknoen, M.DDEF 14A2026-03-260001339970-26-000021Section “Director Compensation”. As printed: 167,422. Filing ↗machine: t19 · r8 · c11 · span 1248898–1248905
12Section 08, Eric BenevichDEF 14A2026-03-260001339970-26-000021Section “Director Compensation”. As printed: 163,849. Filing ↗machine: t19 · r3 · c11 · span 1224148–1224155
13Section 08, Paul Schimmel, Ph.DDEF 14A2026-03-260001339970-26-000021Section “Director Compensation”. As printed: 152,085. Filing ↗machine: t19 · r7 · c11 · span 1243948–1243955
14Section 08, John K. ClarkeDEF 14A2026-03-260001339970-26-000021Section “Director Compensation”. As printed: 19,775. Filing ↗machine: t19 · r4 · c3 · span 1226202–1226208
15Section 08, Sanjay S. Shukla, M.D., M.SDEF 14A2026-03-260001339970-26-000021Section “Director Compensation”. As printed: 0. Filing ↗
16Section 02, Svetlana Lucas 2026-05-11Form 42026-05-130001781292-26-000003Insider equity transaction reported since the proxy statement. Filing ↗
17Section 02, Sara Zaknoen 2026-05-11Form 42026-05-130001602225-26-000003Insider equity transaction reported since the proxy statement. Filing ↗
18Section 02, Paul Schimmel 2026-05-11Form 42026-05-130001236447-26-000003Insider equity transaction reported since the proxy statement. Filing ↗
19Section 02, Eric Benevich 2026-05-11Form 42026-05-130001643653-26-000014Insider equity transaction reported since the proxy statement. Filing ↗
20Section 02, Timothy P. Coughlin 2026-05-11Form 42026-05-130001376043-26-000003Insider equity transaction reported since the proxy statement. Filing ↗
21Section 02, Jane A. Gross 2026-05-11Form 42026-05-130001684258-26-000003Insider equity transaction reported since the proxy statement. Filing ↗
22Section 02, Paul Schimmel 2026-05-27Form 42026-05-280001236447-26-000006Insider equity transaction reported since the proxy statement. Filing ↗
23Section 06, 2025 voteForm 8-K2025-05-060000950170-25-064413Item 5.07 · say-on-pay result. As reported: 97.61% (votes for as printed: 55,237,719). Filing ↗
24Section 06, 2024 voteForm 8-K2024-05-290000950170-24-065899Item 5.07 · say-on-pay result. As reported: 99.07% (votes for as printed: 42,468,397). Filing ↗
25Section 06, 2023 voteForm 8-K2023-05-190000950170-23-023257Item 5.07 · say-on-pay result. As reported: 97.37% (votes for as printed: 37,246,069). Filing ↗
26Section 01 trend, Δ column (computed)ComputedcomputedΔ = current-FY total − prior-FY total per named officer.machine: formula: Δ = FY(n) − FY(n−1)
27Section 03b vehicle mix (computed)ComputedcomputedRSU / PSU / Options grant-date fair values summed per officer from the grants table; % performance-vested = PSU ÷ (RSU+PSU+Options).machine: formula: sums + PSU ratio
28Section 09b board cost (computed)ComputedcomputedCash 370,458 + equity 0 + all other 0 = 370,458, summed across the director compensation table rows.machine: formula: Σ = 370,458
29Scholar Rock Holding Corporation, Section 10b CEO totalDEF 14A2026-04-220001104659-26-046917CEO total compensation as printed in Scholar Rock Holding Corporation's own proxy. As printed: 33,300,927. Filing ↗machine: r1 · c47 · span 363867–363877
30Altimmune, Inc., Section 10b CEO totalDEF 14A2026-03-170001104659-26-028953CEO total compensation as printed in Altimmune, Inc.'s own proxy. As printed: 8,026,590. Filing ↗machine: t81 · r6 · c16 · span 540748–540757
31KalVista Pharmaceuticals, Inc., Section 10b CEO totalDEF 14A2025-08-210001193125-25-185353CEO total compensation as printed in KalVista Pharmaceuticals, Inc.'s own proxy. As printed: 6,772,197. Filing ↗machine: span 291290–291299
32Allakos Inc., Section 10b CEO totalDEF 14A2024-04-230000950170-24-047043CEO total compensation as printed in Allakos Inc.'s own proxy. As printed: 6,299,426. Filing ↗machine: r2 · c26 · span 505098–505107
33Pliant Therapeutics, Inc., Section 10b CEO totalDEF 14A2026-04-220001746473-26-000076CEO total compensation as printed in Pliant Therapeutics, Inc.'s own proxy. As printed: 5,462,901. Filing ↗machine: r2 · c48 · span 436933–436942
34Savara Inc., Section 10b CEO totalDEF 14A2026-04-240001193125-26-177353CEO total compensation as printed in Savara Inc.'s own proxy. As printed: 4,845,000. Filing ↗machine: r2 · c30 · span 340387–340396
35aTYR PHARMA (subject), Section 10b CEO totalDEF 14A2026-03-260001339970-26-000021CEO total compensation as printed in aTYR PHARMA's own proxy. As printed: 3,456,253. Filing ↗machine: t11 · r2 · c21 · span 533754–533763
36X4 Pharmaceuticals, Inc., Section 10b CEO totalDEF 14A2026-03-200001501697-26-000031CEO total compensation as printed in X4 Pharmaceuticals, Inc.'s own proxy. As printed: 2,640,569. Filing ↗machine: span 520865–520874
37Gossamer Bio, Inc., Section 10b CEO totalDEF 14A2026-04-240001728117-26-000027CEO total compensation as printed in Gossamer Bio, Inc.'s own proxy. As printed: 2,300,455. Filing ↗machine: r2 · c48 · span 378559–378568
38Longboard Pharmaceuticals, Inc., Section 10b CEO totalDEF 14A2024-04-120000950170-24-044023CEO total compensation as printed in Longboard Pharmaceuticals, Inc.'s own proxy. As printed: 2,053,804. Filing ↗machine: r2 · c22 · span 476646–476655
39Alector, Inc., Section 10b CEO totalDEF 14A2026-04-280001193125-26-186928CEO total compensation as printed in Alector, Inc.'s own proxy. As printed: 1,995,018. Filing ↗machine: span 1038665–1038674
40Section 10a, Pliant Therapeutics, Inc. reciprocityDEF 14A0001746473-26-000076Pliant Therapeutics, Inc.'s most recent proxy was read; its disclosed peer group does not include aTYR PHARMA. Filing ↗
41Section 10a, Scholar Rock Holding Corporation reciprocityDEF 14A0001104659-26-046917Scholar Rock Holding Corporation's most recent proxy was read; its disclosed peer group does not include aTYR PHARMA. Filing ↗
42Section 10a, Altimmune, Inc. reciprocityDEF 14A0001104659-26-028953Altimmune, Inc. names aTYR PHARMA in its own disclosed peer group (read from its most recent proxy). Filing ↗
43Section 10a, Gossamer Bio, Inc. reciprocityDEF 14A0001728117-26-000027Gossamer Bio, Inc. names aTYR PHARMA in its own disclosed peer group (read from its most recent proxy). Filing ↗
44Section 10a, Savara Inc. reciprocityDEF 14A0001193125-26-177353Savara Inc.'s most recent proxy was read; its disclosed peer group does not include aTYR PHARMA. Filing ↗
45Section 10a, Alector, Inc. reciprocityDEF 14A0001193125-26-186928Alector, Inc.'s most recent proxy was read; its disclosed peer group does not include aTYR PHARMA. Filing ↗
46Section 10a, Allakos Inc. reciprocityDEF 14A0000950170-24-047043Allakos Inc.'s most recent proxy was read; its disclosed peer group does not include aTYR PHARMA. Filing ↗