Velarion · Company Intelligence
Compensation Brief
Fiscal year 2025 · Filings through 2026-10-01

aTYR PHARMA (ATYR)

CEO Sanjay Shukla was paid $3,456,253 in fiscal year 2025, ↑ up 133.4% from fiscal year 2024, the 36th percentile of its disclosed peer group (SEC filing 0001339970-26-000021 ↗): executive and director compensation, incentive-plan design, equity activity, and governance — every figure in this brief is traceable to its exact location in the company’s SEC filing — the document, the section, the page, and the table, linked directly.

$3,456,253 ↑
Sanjay Shukla total, FY2025
36th
CEO pay percentile vs peers
—
CEO-to-median pay ratio
97.61%
Say-on-pay support, latest vote

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Sample answer — “What is the CEO’s total compensation?”
- Sanjay Shukla (President and CEO), FY2025 total compensation $3,456,253 (base salary $638,977) [exec_comp, SEC accession 0001339970-26-000021, filed 2026-03-26]

Answered within 24 hours, every figure cited to its SEC filing. If our database cannot answer your question, you are refunded automatically.

ATYR's next annual pay disclosure, estimated March 26, 2027.est. Filing ↗

01

Summary Compensation Table — fiscal year 2025

Named executive officerSalaryBonusStock awardsOption awardsNon-equity incentivePension / NQDCAll otherTotal
Sanjay Shukla$638,977$0$0$2,595,200$210,862$0$11,214$3,456,253
President and CEO
Jill Broadfoot471,79000713,680113,229011,2141,309,913
Chief Financial Officer
Nancy Denyes451,57000713,680108,377011,2141,284,841
General Counsel

A dash means the company reported no figure in that column; 0 means it reported zero. Figures are as printed in the filing, without adjustment.

Three-year trend — total compensation per named executive

Named executive officerFY2025 totalFY2024 totalFY2023 totalΔ FY2024→2025
Sanjay Shukla$3,456,253$1,480,522$1,300,525↑ +1,975,731
Jill Broadfoot1,309,913746,163677,837↑ +563,750
Nancy Denyes1,284,841729,305651,283↑ +555,536

Prior-year totals as disclosed in the corresponding proxy; deltas are computed values with the derivation shown in the register.

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Companies that benchmark against ATYR
5 companies name ATYR as a compensation peer

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01c

Board of directors

One page per director: the seat, the committees and the fee table, read for one person at a time.

DirectorRoleFY2025 total
Eric BenevichIndependent Director$163,849
Jane A. Gross, Ph.DIndependent Director$170,423
John K. ClarkeFormer director (through FY2025)$19,775
Paul Schimmel, Ph.DIndependent Director$152,085
Sara L. Zaknoen, M.DIndependent Director$167,422
Svetlana Lucas, Ph.DIndependent Director$175,129
Timothy P. CoughlinChair of the Board$194,285
02

Equity transactions since the proxy statement was filed

Every insider equity transaction reported on Form 4 after the 2025 proxy statement — acquisitions, dispositions, exercises, and withholdings — and therefore not reflected in any table in that document.

InsiderRoleTransaction dateCodeSharesPriceValueSecuritySource
Paul SchimmelDirector2026-05-11A50,000$0$0Derivative, Class A CommonForm 4 ↗
Sara ZaknoenDirector2026-05-11A50,00000Derivative, Class A CommonForm 4 ↗
Svetlana LucasDirector2026-05-11A50,00000Derivative, Class A CommonForm 4 ↗
Jane A. GrossDirector2026-05-11A50,00000Derivative, Class A CommonForm 4 ↗
Eric BenevichDirector2026-05-11A50,00000Derivative, Class A CommonForm 4 ↗
Timothy P. CoughlinDirector2026-05-11A50,00000Derivative, Class A CommonForm 4 ↗
Paul SchimmelDirector2026-05-27A100,000049,630CommonForm 4 ↗
Brandon YarasChief Financial Officer2026-10-01A410,00000Derivative, Class A CommonForm 4 ↗

Transaction codes per Form 4: A acquisition · S open-market sale · F tax withholding · M option exercise · G gift. Value = shares × price as reported. Dividend-equivalent and fractional-share accrual rows (derivative Class A, under $10,000 and under 2,000 shares) are aggregated per date; every row appears in full below.

03

Grants of plan-based awards — fiscal year 2025

ExecutiveAwardGrant dateThresholdTargetGrantedMaximumGrant-date fair value
Sanjay ShuklaOption—————$2,595,200
Nancy DenyesOption—————713,680
Jill BroadfootOption—————713,680
03b

Vehicle mix — grant-date fair value by award type

Named executive officerTime-vested (RSU)Performance-vested (PSU)Options / SARs% performance-vested
Sanjay Shukla$0$0$2,595,2000.0%
Nancy Denyes00713,6800.0%
Jill Broadfoot00713,6800.0%

% performance-vested = PSU grant-date fair value ÷ (RSU + PSU + Options), a computed value with the derivation shown in the register.

04

Outstanding equity awards at fiscal year-end 2025

ExecutiveAwardShares unvestedExercise priceMarket value unvested
Sanjay ShuklaOption3,4585.52—
Sanjay ShuklaRSU10,375—8,125
Jill BroadfootOption211,9803.74—
Jill BroadfootRSU3,750—2,937
Nancy DenyesRSU2,687—2,104
Sanjay ShuklaOption53,7792.81—
Sanjay ShuklaOption98,0282.2—
Sanjay ShuklaOption312,5001.5—
Sanjay ShuklaOption770,8343.74—
Jill BroadfootOption1,2505.52—
Jill BroadfootOption7,8492.81—
Jill BroadfootOption24,5072.2—
Jill BroadfootOption78,1251.5—
Nancy DenyesOption8965.52—
Nancy DenyesOption9,1772.81—
Nancy DenyesOption24,5072.2—
Nancy DenyesOption78,1251.5—
Nancy DenyesOption211,9803.74—
05

Incentive-plan design — the Compensation Discussion & Analysis

The Company's executive compensation philosophy is based on aligning executive interests with stockholder interests through long-term incentives linked to Company performance, attracting and retaining superior executive talent, and providing incentives that reward the achievement of performance goals that directly correlate to the enhancement of stockholder value.

CEO STI target (% of salary)55%
CEO LTI target (% of salary)—
Independent consultantRadford
Clawback policytrue
Anti-hedgingtrue
Anti-pledgingfalse
06

Say-on-pay — recent advisory votes

Meeting yearMeeting dateForAgainstAbstainSupportOutcome
20252025-05-0155,237,7191,353,536256,53597.61%Passed
20242024-05-2242,468,397397,92650,83399.07%Passed
20232023-05-1737,246,0691,007,21245,49797.37%Passed
20222022-04-2618,771,20251,25432,56399.73%Passed
20212021-04-287,519,078136,22818,01198.22%Passed

Rows are labeled by annual-meeting year, with the meeting date beside each; the vote held in a given year acts on the prior fiscal year's compensation.

Say-on-pay — history, company's computation

Meeting yearMeeting dateApproval — company's computation
20252025-05-0197.61%a
20242024-05-2299.07%b
20232023-05-1797.37%c

a. Approved under the standard stated in the filing — “Broker non-votes and abstentions will not be counted as votes properly cast with respect to such matter, and will have no effect on the proposal.”.

b. Approved under the standard stated in the filing — “Broker non-votes and abstentions will not be counted as votes properly cast with respect to such matter, and will have no effect on the proposal.”.

c. Approved under the standard stated in the filing — “Broker non-votes and abstentions will not be counted as votes properly cast with respect to such matter, and will have no effect on the proposal.”.

Approval percentages are the company's own reported computations under its own stated standard, not recomputed by Velarion. Where a company does not state a standard, none is assumed.

07

CEO pay ratio

Ratio (CEO : median employee)—
CEO total compensation—
Median employee compensation—
08

Director compensation — fiscal year 2025

DirectorCash feesStock awardsOption awardsNon-equity incentiveAll otherTotal
Timothy P. CoughlinIndependent · Board Chair · Audit$82,200$0$112,085—$0$194,285
Svetlana Lucas, Ph.DIndependent · Audit (Chair) · Partial-year service63,0440112,085—0175,129
Jane A. Gross, Ph.DIndependent · Compensation · Partial-year service58,3380112,085—0170,423
Sara L. Zaknoen, M.DIndependent · Compensation · Partial-year service55,3370112,085—0167,422
Eric BenevichIndependent · Audit · Compensation51,7640112,085—0163,849
Paul Schimmel, Ph.DIndependent40,0000112,085—0152,085
John K. ClarkeIndependent · Not standing for re-election · Departed during FY19,77500—019,775
Sanjay S. Shukla, M.D., M.SEmployee director · Partial-year service000—00
09

Board fee structure and ownership guideline

Annual cash retainer$40,000
Annual equity retainer—
Per board meeting fee—
Committee chair fees (audit / comp / nom-gov)20,000 / 15,000 / 10,000
Stock ownership guideline—
09b

Total cost of the board

ComponentFY2025
Cash retainers and fees$370,458
Equity awards (grant-date value)$0
All other compensation$0
Total cost of the board$370,458

Computed as the sum of the director compensation table rows above; inputs and formula shown in the register.

10

Disclosed peer group — fiscal year 2025

The compensation peer group as the company disclosed it in its own proxy — the reference set behind the percentile above.

10a

Peer reciprocity

Of the companies aTYR PHARMA names as compensation peers, which name it back in their own disclosed peer groups. A peer set the market reciprocates reads differently from one selected in a single direction.

Reciprocity shown where the peer's own current proxy has been read — each mark is cited in the register to that peer's filing. A dash means the peer's current proxy was read and names no reciprocal group including aTYR PHARMA.

10b

Chief executive pay against the peer group

Most recent fiscal-year chief executive compensation at each disclosed peer, taken from that company's own proxy statement.

RankCompanyFYChief executiveSalaryStock awardsOption awardsNon-equity incentiveTotal
1Scholar Rock Holding Corporation2025David Hallal$637,500$22,518,500$7,624,988$452,293$33,300,927
2Altimmune, Inc.2025Vipin Garg658,6941,153,6004,812,167357,7538,026,590
3Allakos Inc.2023Robert Alexander805,4602,171,3402,623,519684,6416,299,426
4Pliant Therapeutics, Inc.2025Bernard Coulie682,0001,810,2502,713,750204,6005,462,901
5KalVista Pharmaceuticals, Inc.2025Benjamin Palleiko477,2023,731,160005,094,999
6Savara Inc.2025Matthew Pauls660,0003,870,0000297,0004,845,000
7Trevi Therapeutics, Inc.2025Jennifer Good622,38202,968,26003,944,892
8aTYR PHARMA2025Sanjay Shukla638,97702,595,200210,8623,456,253
9X4 Pharmaceuticals, Inc.2025Paula Ragan420,1701,081,575002,640,569
10Gossamer Bio, Inc.2025Faheem Hasnain649,64901,392,006254,8002,300,455
11Longboard Pharmaceuticals, Inc.2023Kevin Lind610,0000986,934442,8602,053,804
12Alector, Inc.2025Arnon Rosenthal678,100422,876558,216325,4881,995,018
13Kezar Life Sciences, Inc.2024Christopher Kirk600,00000313,500927,300
—Acelyrin, Inc.——————not in coverage universe
—Fortress Biotech, Inc.——————not in coverage universe
—Immunic, Inc.——————not in coverage universe
—Jasper Therapeutics, Inc.——————not in coverage universe

Each peer's compensation is from its most recently filed proxy; fiscal years differ by company. Each peer's figures come from its own filings; the subject company is highlighted. Peers whose chief executive compensation is outside our coverage universe are stated as such, never left blank.

10c

Incentive design against the same peers

CompanyAnnual incentive payoutLong-term vehicle mixPSU measures include TSRCompensation consultant
aTYR PHARMA60% (Committee lowered payout from 75% achievement to 60% given EFZO-FIT study results)%Option 100not disclosedRadford
Acelyrin, Inc.not disclosednot disclosednot disclosednot disclosed
Alector, Inc.80%not disclosednot disclosedPearl Meyer & Partners, LLC
Allakos Inc.not disclosedOptions 55 · RSU 45NoCompensia
Altimmune, Inc.not disclosednot disclosednot disclosednot disclosed
Fortress Biotech, Inc.not disclosednot disclosednot disclosednot disclosed
Gossamer Bio, Inc.65%not disclosedNoAlpine Rewards
Immunic, Inc.not disclosednot disclosednot disclosednot disclosed
Jasper Therapeutics, Inc.not disclosednot disclosednot disclosednot disclosed
KalVista Pharmaceuticals, Inc.not disclosednot disclosedNoAon
Kezar Life Sciences, Inc.not disclosednot disclosednot disclosednot disclosed
Longboard Pharmaceuticals, Inc.not disclosedOption 100not disclosedFrederic W. Cook & Co.
Pliant Therapeutics, Inc.not disclosedOptions 67 · RSU 33not disclosedAlpine Rewards
Savara Inc.90%RSU 100not disclosedAon
Scholar Rock Holding Corporationnot disclosednot disclosedNoPay Governance
Trevi Therapeutics, Inc.not disclosednot disclosednot disclosednot disclosed
X4 Pharmaceuticals, Inc.not disclosednot disclosednot disclosednot disclosed

Read from each company's Compensation Discussion and Analysis: annual incentive payout as a percentage of target, the disclosed long-term vehicle mix, whether performance-share measures include a total shareholder return component, and the retained compensation consultant. “Not disclosed” appears only where the company's filing does not state the item.

10d

Frequently asked questions

Who was the highest-paid executive at aTYR PHARMA in fiscal 2025?
As disclosed, Sanjay Shukla (President and CEO) was the highest-paid executive at aTYR PHARMA in fiscal 2025, with total compensation of $3,456,253.
How does aTYR PHARMA's chief executive pay compare to its disclosed peer group?
As disclosed, Sanjay Shukla's total compensation ranked at the 36th percentile of aTYR PHARMA's disclosed peer group.
How did shareholders vote on aTYR PHARMA's executive compensation?
As disclosed, 97.61% of shareholders supported aTYR PHARMA's executive compensation in the latest say-on-pay vote.
How much does aTYR PHARMA spend in total on board compensation for fiscal 2025?
As disclosed, aTYR PHARMA reported total board compensation of $370,458 for fiscal 2025.

Answers are drawn only from the figures disclosed in this brief and cited to their location in the source filing. Where a figure required for an answer is not disclosed, that is stated rather than estimated.

11

Source register

Each figure's location is given both for the eye (section · page · table · row · column) and for a script (table/row/column index and exact character span in the filed document). Any figure can be independently confirmed against the source.

#SupportsDocumentFiledAccessionLocation in document
1Section 01, Sanjay ShuklaDEF 14A2026-03-260001339970-26-000021Section “Executive Compensation”. As printed: 3,456,253. Filing ↗machine: r2 · c21 · span 533754–533763
2Section 01, Jill BroadfootDEF 14A2026-03-260001339970-26-000021Section “Executive Compensation”. As printed: 1,309,913. Filing ↗machine: r4 · c21 · span 549228–549237
3Section 01, Nancy DenyesDEF 14A2026-03-260001339970-26-000021Section “Executive Compensation”. As printed: 1,284,841. Filing ↗machine: r6 · c21 · span 564707–564716
4Section 03, Sanjay Shukla OptionDEF 14A2026-03-260001339970-26-000021Section “SUMMARY COMPENSATION TABLE” · grants of plan-based awards table. Filing ↗machine: t11 · r2 · c9 · span 529802–529811
5Section 03, Nancy Denyes OptionDEF 14A2026-03-260001339970-26-000021Section “SUMMARY COMPENSATION TABLE” · grants of plan-based awards table. Filing ↗machine: t11 · r6 · c9 · span 560757–560764
6Section 03, Jill Broadfoot OptionDEF 14A2026-03-260001339970-26-000021Section “SUMMARY COMPENSATION TABLE” · grants of plan-based awards table. Filing ↗machine: t11 · r4 · c9 · span 545278–545285
7Section 07, pay ratioDEF 14A2026-03-260001339970-26-000021Section “CEO Pay Ratio”. As printed: —. Filing ↗
8Section 08, Timothy P. CoughlinDEF 14A2026-03-260001339970-26-000021Section “Director Compensation”. As printed: 194,285. Filing ↗
9Section 08, Svetlana Lucas, Ph.DDEF 14A2026-03-260001339970-26-000021Section “Director Compensation”. As printed: 175,129. Filing ↗machine: span 1238808–1239017
10Section 08, Jane A. Gross, Ph.DDEF 14A2026-03-260001339970-26-000021Section “Director Compensation”. As printed: 170,423. Filing ↗machine: span 1233858–1234067
11Section 08, Sara L. Zaknoen, M.DDEF 14A2026-03-260001339970-26-000021Section “Director Compensation”. As printed: 167,422. Filing ↗machine: span 1248707–1248916
12Section 08, Eric BenevichDEF 14A2026-03-260001339970-26-000021Section “Director Compensation”. As printed: 163,849. Filing ↗machine: span 1223957–1224166
13Section 08, Paul Schimmel, Ph.DDEF 14A2026-03-260001339970-26-000021Section “Director Compensation”. As printed: 152,085. Filing ↗machine: span 1243757–1243966
14Section 08, John K. ClarkeDEF 14A2026-03-260001339970-26-000021Section “Director Compensation”. As printed: 19,775. Filing ↗machine: span 1226011–1226219
15Section 08, Sanjay S. Shukla, M.D., M.SDEF 14A2026-03-260001339970-26-000021Section “Director Compensation”. As printed: 0. Filing ↗
16Section 02, Paul Schimmel 2026-05-11Form 42026-05-130001236447-26-000003Insider equity transaction reported since the proxy statement. Filing ↗
17Section 02, Sara Zaknoen 2026-05-11Form 42026-05-130001602225-26-000003Insider equity transaction reported since the proxy statement. Filing ↗
18Section 02, Svetlana Lucas 2026-05-11Form 42026-05-130001781292-26-000003Insider equity transaction reported since the proxy statement. Filing ↗
19Section 02, Jane A. Gross 2026-05-11Form 42026-05-130001684258-26-000003Insider equity transaction reported since the proxy statement. Filing ↗
20Section 02, Eric Benevich 2026-05-11Form 42026-05-130001643653-26-000014Insider equity transaction reported since the proxy statement. Filing ↗
21Section 02, Timothy P. Coughlin 2026-05-11Form 42026-05-130001376043-26-000003Insider equity transaction reported since the proxy statement. Filing ↗
22Section 02, Paul Schimmel 2026-05-27Form 42026-05-280001236447-26-000006Insider equity transaction reported since the proxy statement. Filing ↗
23Section 02, Brandon Yaras 2026-10-01Form 42026-10-010002156611-26-000010Insider equity transaction reported since the proxy statement. Filing ↗
24Section 06, 2025 voteForm 8-K2025-05-060000950170-25-064413Item 5.07 · say-on-pay result. As reported: 97.61% (votes for as printed: 55,237,719). Filing ↗
25Section 06, 2024 voteForm 8-K2024-05-290000950170-24-065899Item 5.07 · say-on-pay result. As reported: 99.07% (votes for as printed: 42,468,397). Filing ↗
26Section 06, 2023 voteForm 8-K2023-05-190000950170-23-023257Item 5.07 · say-on-pay result. As reported: 97.37% (votes for as printed: 37,246,069). Filing ↗
27Section 01 trend, Δ column (computed)Computed——Δ = current-FY total − prior-FY total per named officer.machine: formula: Δ = FY(n) − FY(n−1)
28Section 03b vehicle mix (computed)Computed——RSU / PSU / Options grant-date fair values summed per officer from the grants table; % performance-vested = PSU ÷ (RSU+PSU+Options).machine: formula: sums + PSU ratio
29Section 09b board cost (computed)Computed——Cash 370,458 + equity 0 + all other 0 = 370,458, summed across the director compensation table rows.machine: formula: Σ = 370,458
30Scholar Rock Holding Corporation, Section 10b CEO totalDEF 14A2026-04-220001104659-26-046917CEO total compensation as printed in Scholar Rock Holding Corporation's own proxy. As printed: 33,300,927. Filing ↗machine: r1 · c47 · span 363867–363877
31Altimmune, Inc., Section 10b CEO totalDEF 14A2026-03-170001104659-26-028953CEO total compensation as printed in Altimmune, Inc.'s own proxy. As printed: 8,026,590. Filing ↗
32Allakos Inc., Section 10b CEO totalDEF 14A2024-04-230000950170-24-047043CEO total compensation as printed in Allakos Inc.'s own proxy. As printed: 6,299,426. Filing ↗machine: r2 · c26 · span 505098–505107
33Pliant Therapeutics, Inc., Section 10b CEO totalDEF 14A2026-04-220001746473-26-000076CEO total compensation as printed in Pliant Therapeutics, Inc.'s own proxy. As printed: 5,462,901. Filing ↗machine: r2 · c48 · span 436933–436942
34KalVista Pharmaceuticals, Inc., Section 10b CEO total10-K/A2026-04-300001193125-26-197556CEO total compensation as printed in KalVista Pharmaceuticals, Inc.'s own proxy. As printed: 5,094,999. Filing ↗
35Savara Inc., Section 10b CEO totalDEF 14A2026-04-240001193125-26-177353CEO total compensation as printed in Savara Inc.'s own proxy. As printed: 4,845,000. Filing ↗machine: r2 · c30 · span 340387–340396
36Trevi Therapeutics, Inc., Section 10b CEO totalDEF 14A2026-04-210001193125-26-166695CEO total compensation as printed in Trevi Therapeutics, Inc.'s own proxy. As printed: 3,944,892. Filing ↗machine: r2 · c21 · span 594331–594340
37aTYR PHARMA (subject), Section 10b CEO totalDEF 14A2026-03-260001339970-26-000021CEO total compensation as printed in aTYR PHARMA's own proxy. As printed: 3,456,253. Filing ↗machine: r2 · c21 · span 533754–533763
38X4 Pharmaceuticals, Inc., Section 10b CEO totalDEF 14A2026-03-200001501697-26-000031CEO total compensation as printed in X4 Pharmaceuticals, Inc.'s own proxy. As printed: 2,640,569. Filing ↗machine: span 520865–520874
39Gossamer Bio, Inc., Section 10b CEO totalDEF 14A2026-04-240001728117-26-000027CEO total compensation as printed in Gossamer Bio, Inc.'s own proxy. As printed: 2,300,455. Filing ↗machine: r2 · c48 · span 378559–378568
40Longboard Pharmaceuticals, Inc., Section 10b CEO totalDEF 14A2024-04-120000950170-24-044023CEO total compensation as printed in Longboard Pharmaceuticals, Inc.'s own proxy. As printed: 2,053,804. Filing ↗machine: r2 · c22 · span 476646–476655
41Alector, Inc., Section 10b CEO totalDEF 14A2026-04-280001193125-26-186928CEO total compensation as printed in Alector, Inc.'s own proxy. As printed: 1,995,018. Filing ↗machine: span 1038665–1038674
42Kezar Life Sciences, Inc., Section 10b CEO totalDEF 14A2025-04-250001645666-25-000020CEO total compensation as printed in Kezar Life Sciences, Inc.'s own proxy. As printed: 927,300. Filing ↗machine: span 286147–286154
43Section 10a, Pliant Therapeutics, Inc. reciprocityDEF 14A—0001746473-26-000076Pliant Therapeutics, Inc.'s most recent proxy was read; its disclosed peer group does not include aTYR PHARMA. Filing ↗
44Section 10a, Scholar Rock Holding Corporation reciprocityDEF 14A—0001104659-26-046917Scholar Rock Holding Corporation's most recent proxy was read; its disclosed peer group does not include aTYR PHARMA. Filing ↗
45Section 10a, Allakos Inc. reciprocityDEF 14A—0000950170-24-047043Allakos Inc.'s most recent proxy was read; its disclosed peer group does not include aTYR PHARMA. Filing ↗
46Section 10a, Altimmune, Inc. reciprocityDEF 14A—0001104659-26-028953Altimmune, Inc. names aTYR PHARMA in its own disclosed peer group (read from its most recent proxy). Filing ↗
47Section 10a, Savara Inc. reciprocityDEF 14A—0001193125-26-177353Savara Inc.'s most recent proxy was read; its disclosed peer group does not include aTYR PHARMA. Filing ↗
48Section 10a, Alector, Inc. reciprocityDEF 14A—0001193125-26-186928Alector, Inc.'s most recent proxy was read; its disclosed peer group does not include aTYR PHARMA. Filing ↗
49Section 10a, Gossamer Bio, Inc. reciprocityDEF 14A—0001728117-26-000027Gossamer Bio, Inc. names aTYR PHARMA in its own disclosed peer group (read from its most recent proxy). Filing ↗
50Section 10a, Trevi Therapeutics, Inc. reciprocityDEF 14A—0001193125-26-166695Trevi Therapeutics, Inc.'s most recent proxy was read; its disclosed peer group does not include aTYR PHARMA. Filing ↗

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